ABSTRACT Here we describe two new notoungulate taxa from early Oligocene deposits of the Abanico Formation in the eastern Tinguiririca valley of the Andes of central Chile, including a notostylopid (gen. et sp. nov.) and three basal toxodontians, cf. Homalodotheriidae, one of which is formally named a new species. The valley's eponymous fossil mammal fauna became the basis for recognizing a new South American Land Mammal “Age” intervening between the Mustersan and Deseadan of the classical SALMA sequence, the Tinguirirican. As a temporal intermediate between the bracketing SALMAs (Deseadan and Mustersan), the Tinguirirican is characterized by a unique cooccurrence of taxa otherwise known either from demonstrably younger or more ancient deposits, as well as some taxa with temporal ranges restricted to this SALMA. In this regard, two of the notoungulates described here make their last known stratigraphic appearances in the Tinguiririca Fauna, Chilestylops davidsoni (gen. et sp. nov.), the youngest notostylo...

We describe a new species of scaphitid ammonite from the Upper Cretaceous (lower Maastrichtian) of North America. Hoploscaphites sargklofak, n. sp., is endemic to the U.S. Western Interior, but closely resembles H. constrictus Sowerby, 1817, from the Maastrichtian of northern Europe.

ABSTRACT A new genus, Hexapopha, is described for a group of species from Costa Rica that resemble those of Coxapopha Platnick in having an elevated carapace and heavily sclerotized, highly modified male endites but differ in retaining six eyes, lacking leg spines, and having no female genitalic scape. Xestaspis reimoseri Fage is transferred to Hexapopha as the type species, its female is described for the first time, and three new species (H. hone, H. jimenez, and H. osa) are described. Dysderina caeca Biraben, from Argentina, is transferred to Coxapopha.

The Costarina faunas of Panama and Colombia are revised. Because Chickering described the sexes from his Panamanian collections separately, several of his names based only on females are placed as junior synonyms of names based only on males: C. humphreyi (Chickering) is newly synonymized with C. recondita (Chickering), as are C. belinda (Chickering) with C. intempina (Chickering), C. rigida (Chickering) with C. abdita (Chickering), C. silvatica (Chickering) with C. dura (Chickering), and C. improvisa (Chickering) with C. seclusa (Chickering). The Costa Rican species C. pittier Platnick and Berniker is newly recorded from Panama. Nine new species are described from Panama (C. sorkini, cerrocol, bocas, chiriqui, clara, fortuna, almirante, dybasi, and tskui), as are 11 from Colombia (C. sasaima, suiza, otun, choco, yotoco, saladito, antonio, anchicaya, gorgona, helechal, and taraira). Previous studies on more northern species are supplemented; newly available collections from Nicaragua reveal the presence there of two new species (C. kilambe and diablo), and locality records are provided for the Costa Rican members of the genus.

Scorpions of the genus Heterometrus Ehrenberg, 1828, are distributed from India and Sri Lanka throughout the Southeast Asian mainland and archipelagos as far as Wallaces Line. Despite this widespread distribution, Heterometrus was not recorded from Pakistan until a single specimen from Azad Kashmir was reported from the collection of the Pakistan Museum of Natural History, Islamabad. Perhaps because the specimen was misidentified as Heterometrus wroughtoni (Pocock, 1899), a species that occurs much farther to the southeast in India, the presence of Heterometrus in Pakistan remained uncertain until fresh material of a distinctive species of Heterometrus was recently collected at several locations in Khyber Pakhtoon Khawa. After comparison of the material with the holotype and only known specimen of a little-known species, Heterometrus latimanus (Pocock, 1894), with an indefinite type locality in “India,” the Pakistani material was determined to be conspecific. In the present contribution, H. latimanus is redescribed based on adult males and females from several localities, and the specimen from Azad Kashmir tentatively assigned to it, confirming the presence of Heterometrus in Pakistan. The new locality records extend the distribution of the genus considerably to the northwest, and west of the Indus River for the first time. The known records of H. latimanus appear to be isolated from other Heterometrus occurring on the Indian subcontinent by the Great Indian (Thar) Desert, an arid, sandy basin extending from eastern Pakistan to northwestern India.

Age-graded fossils of Pleistocene endemic Cretan deer (Candiacervus spp.) reveal unexpectedly high juvenile mortality similar to that reported for extant mainland ruminants, despite the fact that these deer lived in a predator-free environment and became extinct before any plausible date for human arrival. Age profiles show that deer surviving past the fawn stage were relatively long-lived for ruminants, indicating that high juvenile mortality was not an expression of their living a “fast” life. Although the effects on survivorship of such variables as fatal accidents, starvation, and disease are difficult to gauge in extinct taxa, the presence of extreme morphological variability within nominal species/ecomorphs of Candiacervus is consistent with the view that high juvenile mortality can function as a key innovation permitting rapid adaptation in insular contexts.

ABSTRACT We describe and name three new species of Musseromys from the mountains of northern Luzon based on morphological and DNA sequence data. Previously, Musseromys was known only from one species from the lowlands of central Luzon. These are the smallest-known members of the cloud rat clade of endemic Philippine murids, weighing only 15–22 g, an order of magnitude smaller than the previously smallest known members of the clade (Carpomys spp.), and more than two orders of magnitude smaller than the largest members (Phloeomys spp.). These discoveries raise the number of native murids documented on Luzon to 43, 93% of which are endemic, and 88% of which are members of two endemic Philippine clades. Musseromys is inferred to have originated in montane habitats, probably in the Central Cordillera of northern Luzon, with movement to two other areas on Luzon, one in montane habitat and one in lowland habitat, associated with the speciation process.

The partially submerged Impossível-Ioiô cave system located in the karst region of Chapada Diamantina in Bahia (Brazil) has recently been the target of extensive paleontological studies. Here we provide the first report of fossil bats from this cave system, in which we recognize six species based on humeral remains: Furipterus horrens, Chrotopterus auritus, Mormoops cf. megalophylla, Pteronotus gymnonotus, Pteronotus parnellii, and Noctilio leporinus. Morphology of the humerus of these taxa is described in a comparative framework to document taxonomic assessments and provide a basis for future studies of fossil bat faunas. The relevance of the new records reported here is evaluated at a broader continental scale, as well as in contrast with the recent bat fauna of the region. The record of Noctilio leporinus stands as the first fossil occurrence of this species on the South American continent.

A new genus, Varioonops, is established for a group of species characterized by sexual dimorphism in abdomen morphology (with a dorsal scutum present in males but not females), male palps with a separate cymbium and bulb, and a patterned abdomen. The closest relatives of the new genus appear to be three similar, but as yet undescribed, Neotropical genera that share those characters, but the members of Varioonops are united by a putative synapomorphy that does not seem to occur in those undescribed groups: a granulated sternum. Attention is drawn to the cymbial cone, a structure (possibly a receptor) found near the tip of the cymbium in Varioonops, members of the Orchestininae, and at least some soft-bodied members of the Oonopinae. A total of 23 new species are described: V. cafista (the type species), V. velsala, V. montesta, V. spatharum, V. poas, V. varablanca, V. tortuguero, V. veragua, V. heredia, and girven from Costa Rica, V. ramila and V. sansidro from Costa Rica and Panama, V. funator and cerrado from Panama, V. edvardi and V. sinesama from Colombia, and V. yacambu, V. trujillo, V. pittieri, V. chordio, V. parlata, V. potaguo, and V. grancho from Venezuela.

We describe Hoploscaphites gilberti, n. sp. (Ammonoidea: Ancyloceratina), from the Upper Cretaceous (middle-upper Campanian) Pierre Shale spanning the zones of Baculites scotti and Didymoceras nebrascense in Colorado, Wyoming, Montana, and South Dakota. This species is strongly dimorphic and is characterized by a compressed whorl section, with a rounded to elongate outline in lateral view. The apertural angle is approximately 50° in macroconchs. The body chamber is ornamented with fine flexuous ribs, umbilicolateral bullae, and ventrolateral tubercles. Hoploscaphites gilberti, n. sp., most closely resembles H. gilli Cobban and Jeletzky, 1965, but differs from this species in several important features: (1) the flanks of the body chamber are nearly subparallel rather than steeply convergent toward the venter, (2) the ventrolateral tubercles are larger and more numerous, and (3) the ribs are more widely spaced. Hoploscaphites gilberti, n. sp., is abundant at “tepee buttes” in the Pierre Shale, which are now interpreted as cold methane seeps.